Die ungleichen Cousins David (Jesse Eisenberg) und Benji (Kieran Culkin) reisen gemeinsam nach Polen, um mehr über das Leben ihrer kürzlich verstorbenen Großmutter zu erfahren. Der Roadtrip, der als Reise in die Vergangenheit gedacht war, konfrontiert die beiden mit der Gegenwart, in der sie ihre Beziehung zueinander und ihre Familiengeschichte hinterfragen.
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